E Chereskin, SJ Allen, RC Connor, M Krützen, SL King (2024). In pop pursuit: social bond strength predicts vocal synchrony during cooperative mate guarding in bottlenose dolphins. Philosophical Transactions B 379 (1905), 20230194. DOI: 10.1098/rstb.2023.0194


Peters KJ, Gerber L, Scheu L, Cicciarella, Zoller JA, Fei Z, Horvath S, Allen SJ, King SL, Connor RC, Rollins LA, Krützen M (2023). An epigenetic DNA methylation clock for age estimates in Indo-Pacific bottlenose dolphins (Tursiops aduncus). Evolutionary Applications 16: 126-133. DOI: 10.1111/eva.13516

Sørensen PM, Haddock A, Guarino E, Jaakkola K, McMullan C, Jensen FH, Tyack PL, King SL (2023). Anthropogenic noise impairs cooperation in bottlenose dolphins. Current Biology  DOI: https://doi.org/10.1016/j.cub.2022.12.063

Wittwer S, Gerber L, Allen SJ, Willems EP, Marfurt SM, Krützen M (2023). Reconstructing the colonization history of Indo-Pacific bottlenose dolphins (Tursiops aduncus) in Northwestern Australia. Molecular Ecology DOI: https://doi.org/10.1111/mec.16984


Brown AM, Allen SJ, Kelly N, Hodgson A (2022). Using Unoccupied Aerial Vehicles to investigate availability and group size error for aerial surveys of coastal dolphins. Remote Sensing in Ecology and Conservation https://doi.org/10.1002/rse2.313.

Chabanne DB, Krützen M, Finn H, Allen SJ (2022). Evidence of male alliance formation within a small dolphin community. Mammalian Biology https://doi.org/10.1007/s42991-022-00295-7.

Chereskin E, Connor RC, Friedman WR, Jensen FH, Allen SJ, Sørensen PM, Krützen M and King SL (2022). Allied male dolphins use vocal exchanges to ‘‘bond at a distance’’. Current Biology https://doi.org/10.1016/j.cub.2022.02.019.

Connor RC, Krützen M, Allen SJ, Sherwin WB, King SL (2022). Strategic intergroup alliances increase access to a contested resource in male bottlenose dolphins. Proceedings of the National Academy of Sciences 119: e2121723119. www.pnas.org/doi/10.1073/pnas.2121723119.

Dolman S, Breen C, Brakes P, Butterworth A and Allen SJ (2022). The individual welfare concerns for small cetaceans from two bycatch mitigation techniques. Marine Policy 143: 105126. https://doi.org/10.1016/j.marpol.2022.105126.

Gerber L, Connor RC, Allen SJ, Horlacher K, King SL, Sherwin WB, Willems E, Wittwer S and Krützen M (2022). Social integration influences fitness in allied male dolphins. Current Biology https://doi.org/10.1016/j.cub.2022.03.027.

Hamilton RA, Gazda SL, King SL, Stakhammar J and Connor RC (2022). Bottlenose dolphin communication during a role-specialized group foraging task. Behavioural Processes https://doi.org/10.1016/j.beproc.2022.104691.

King SL and Jensen FH (2022). Rise of the machines: Integrating technology with playback experiments to study cetacean social cognition in the wild. Methods in Ecology and Evolution https://doi.org/10.1111/2041-210X.13935.

Manlik O, Lacy RC, Sherwin WB, Finn H, Loneragan NR and Allen SJ (2022). A stochastic model for estimating sustainable limits to wildlife mortality in a changing world. Conservation Biology e13897 DOI: 10.1111/cobi.13897.

Marfurt SM, Allen SJ, Bizzozzero MR, Willems EP, King SL, Connor RC, Kopps AM, Wild S, Gerber L, Wittwer S, Krützen M (2022). Association patterns and community structure among female bottlenose dolphins: environmental, genetic and cultural factors. Mammalian Biology https://doi.org/10.1007/s42991-022-00259-x.


Chabanne DBH, Allen SJ, Sherwin WB, Finn H, Krützen M (2021). Inconsistency between socio-spatial and genetic structure in a coastal dolphin population. Frontiers in Marine Science 7: 617540. https://www.frontiersin.org/articles/10.3389/fmars.2020.617540/full.

Gerber L, Wittwer S, Allen SJ, Holmes K, King SL, Wild S, Willems E, Connor RC, Krützen M (2021). Cooperative partner choice in multi-level dolphin alliances. Scientific Reports 11: 6901. https://www.nature.com/articles/s41598-021-85583-x.

King SL, Connor RC, Krützen M, Allen SJ (2021). Cooperation-based concept formation in male bottlenose dolphins. Nature Communications 12: 2373. http://disq.us/t/3wd8916.

King SL, Guarino E, Donegan K, McMullen C, Jaakkola K (2021). Evidence that bottlenose dolphins can communicate with vocal signals to solve a cooperative task. Royal Society of Open Science 8: 202073. https://doi.org/10.1098/rsos.202073.


Gerber L, Connor RC, King SL, Allen SJ, Wittwer S, Bizzozzero M, Friedman W, Kalberer S, Sherwin W, Wild S, Willems E, Krützen M (2020). Affiliation history and age similarity predict alliance formation in adult male bottlenose dolphins. Behavioral Ecology 31: 361-370. https://doi.org/10.1093/beheco/arz195.

Hunt TN, Allen SJ, Bejder L, Parra GJ (2020). Identifying priority habitat for conservation and management of Australian humpback dolphins within a marine protected area. Scientific Reports 10: 14366. https://doi.org/10.1038/s41598-020-69863-6.

Moore BL, Connor RC, Allen SJ, Krützen M, King SL (2020). Acoustic coordination by allied male dolphins in a cooperative context. Proceedings of the Royal Society B 287: 20192944. http://dx.doi.org/10.1098/rspb.2019.2944.

McCue LM, Cioffi WR, Heithaus MR, Barrè L, Connor RC (2020). Synchrony, leadership, and association in male Indo‐Pacific bottlenose dolphins (Tursiops aduncus). Ethology https://onlinelibrary.wiley.com/doi/abs/10.1111/eth.13025.

O’Brien O, Allen SJ, Krützen M, Connor RC (2020). Alliance-specific habitat selection by male Indo-Pacific bottlenose dolphins in Shark Bay, Western Australia. Animal Behaviour 164: 39-49. https://www.sciencedirect.com/science/article/abs/pii/S0003347220300828?via%3Dihub.

Wild S, Hoppitt WJE, Allen SJ, Krützen M (2020). Integrating genetic, environmental and social networks to reveal transmission pathways of a dolphin foraging innovationCurrent Biology 30: 1-7. https://doi.org/10.1016/j.cub.2020.05.069.


Bizzozzero MR, Allen SJ, Gerber L, King SL, Wild S, Connor RC, Friedman W, Wittwer S, Krützen M (2019). Tool use and alliance partner choice in male bottlenose dolphins. Proceedings of the Royal Society B 286: 20190898. http://dx.doi.org/10.1098/rspb.2019.0898

Connor RC, Sakai M, Morisaka T, Allen SJ (2019). The Indo-Pacific bottlenose dolphin Tursiops aduncus. Ch16 in Ethology and Behavioral Ecology of Odontocetes (ed. Würsig). Springer Nature. Pp 345-368. DOI: 10.1007/978-3-030-16663-2.

Hamilton RA, Borcuch T, Allen SJ, Cioffi WR, Bucci V, Krützen M, Connor RC (2019). Aggression varies with consortship rate and habitat in a dolphin social network. Behavioral Ecology & Sociobiology 73: 141. https://doi.org/10.1007/s00265-019-2753-1.

Hunt TN, Allen SJ, Bejder L, Parra GJ (2019). Assortative interactions revealed in a fission–fusion society of Australian humpback dolphins. Behavioral Ecology 29: 1-14. DOI: 10.1093/beheco/arz029.

King SL, Allen SJ, Krützen M, Connor RC (2019). Vocal behaviour of allied male dolphins during cooperative mate guarding. Animal Cognition https://doi.org/10.1007/s10071-019-01290-1.

Manlik O, Chabanne D, Daniel C, Bejder L, Allen SJ, Sherwin WB (2019). Demography and genetics suggest reversal of dolphin source-sink dynamics, with implications for conservation. Marine Mammal Science 35: 732–759. DOI: 10.1111/mms.12555.

Manlik O, Krützen M, Kopps AM, Mann J, Bejder L, Allen SJ, Frère C, Connor RC, Sherwin WB (2019). Is MHC diversity a better marker for conservation than neutral genetic diversity? A case study of two contrasting dolphin populations. Ecology and Evolution 2019;00: 1-13 DOI: 10.1002/ECE3.5265.

Wild S, Allen SJ, Krützen M, King SL, Gerber L, Hoppitt WJE (2019). Multi-network-based diffusion analysis reveals vertical cultural transmission of sponge tool use within dolphin matrilines. Biology Letters 15: 20190227. http://dx.doi.org/10.1098/rsbl.2019.0227.

Wild S, Krützen M, Rankin RW, Hoppitt WJE, Gerber L, Allen SJ (2019). Long-term decline in survival and reproduction of dolphins following a marine heatwave. Current Biology 29: R239-240. DOI: 10.1016/j.cub.2019.02.047.


Jaakkola K, Guarino E, Donegan K, King SL (2018). Bottlenose dolphins can understand their partner’s role in a cooperative task. Proceedings of the Royal Society B 285 https://doi.org/10.1098/rspb.2018.0948.

King SL, Friedman W, Allen SJ, Gerber L, Jensen F, Wittwer S, Connor RC, Krützen M (2018). Bottlenose dolphins retain individual vocal labels in multi-level alliances. Current Biology https://doi.org/10.1016/j.cub.2018.05.013.

Smith F, Allen SJ, Bejder L, Brown AM (2018). Shark bite injuries on three inshore dolphin species in tropical northwestern Australia. Marine Mammal Science 34: 87-99. DOI: 10.1111/mms.12435. http://rdcu.be/vavT.


Allen SJ, King SL, Krützen M, Brown AM (2017). Multi-modal sexual displays in Australian humpback dolphins. Scientific Reports 7: 13644. http://rdcu.be/w3tL.

Allen SJ, Pollock KH, Bouchet PJ, Kobryn HT, McElligott DB, Nicholson KE, Smith JN, Loneragan NR (2017). Preliminary estimates of the abundance and fidelity of dolphins in a demersal trawl fishery. Scientific Reports 7: 4995. http://www.nature.com/articles/s41598-017-05189-0.

Connor RC, Cioffi W, Randić S, Allen SJ, Watson-Capps JJ, Krützen M (2017). Male alliance behaviour and mating access varies with habitat in a dolphin social network. Scientific Reports 7:46354. DOI: 10.1038/srep46354. http://www.nature.com/articles/srep46354.

Hunt TN, Bejder L, Allen SJ, Rankin RW, Hanf D, Parra GJ (2017). Demographic characteristics of Australian humpback dolphins reveal important habitat toward the south-western limit of their range. Endangered Species Research 32: 71-88. https://doi.org/10.3354/esr00784.


Allen SJ, Bryant K, Kraus R, Loneragan N, Kopps A, Brown A, Gerber L, Krützen M (2016). Genetic isolation between coastal and fishery-impacted, offshore bottlenose dolphin (Tursiops spp.) populations. Molecular Ecology DOI: 10.1111/mec.13622.

Brown AM, Bejder L, Parra GJ, Cagnazzi D, Hunt T, Smith JL, Allen SJ (2016). Sexual dimorphism and geographic variation in dorsal fin features of Australian humpback dolphins, Sousa sahulensis. Advances in Marine Biology 73: 273-314.

Brown AM, Bejder L, Pollock KH, Allen SJ (2016). Site-specific assessments of the abundance of three inshore dolphin species to inform conservation and management. Frontiers in Marine Science 3: 4. DOI: 10.3389/fmars.2016.00004.

Higham JES, Bejder L, Allen SJ, Corkeron PJ, Lusseau D (2016). Managing whale-watching as a non-lethal consumptive activity.  Journal of Sustainable Tourism 24(1): 73-90.

Hiley H, Perry S, Hartley S, King SL (2016). What’s Occurring? Ultrasonic signature whistle use in Welsh bottlenose dolphins (Tursiops truncatus). Bioacoustics DOI: 10.1080/09524622.2016.1174885.

Jedensjö M, Kemper CM, Krützen M (2016). Cranial morphology and taxonomic resolution of some dolphin taxa (Delphinidae) in Australian waters, with a focus on the genus TursiopsMarine Mammal Science https://onlinelibrary.wiley.com/journal/10.1111/(ISSN)1748-7692/earlyview.

King SL, Allen SJ, Connor RC, Jaakkola K (2016). Cooperation or dolphin ‘tug-of-war’? Comment on Kuczaj et al. and Eskelinen et al. Animal Cognition DOI 10.1007/s10071-016-1026-x.

King SL, Guarino E, Donegan K, Hecksher J, Jaakkola K (2016). Further insights into postpartum signature whistle use in bottlenose dolphins (Tursiops truncatus). Marine Mammal Science DOI: 10.1111/mms.12317.

King SL, Guarino E, Keaton L, Erb L, Jaakkola K (2016). Maternal signature whistle use aids mother calf reunions in a bottlenose dolphin, Tursiops truncatus. Behavioural Processes 126: 64-70.

King SL, McGregor PM (2016) Vocal Matching: the what, the why and the how. Biology Letters 12(10): 20160666.

Rankin R, Nicholson K, Allen S, Krützen M, Bejder L, Pollock K (2016). A full-capture Hierarchical Bayesian model of Pollock’s closed robust design and application to dolphins. Frontiers in Marine Science 3: 25. DOI: 10.3389/fmars.2016.00025.

Sprogis K, Pollock K, Raudino H, Allen SJ, Kopps AM, Manlik O, Tyne J, Bejder L (2016). Sex-specific patterns of abundance, temporary emigration and survival of Indo-Pacific bottlenose dolphins (Tursiops aduncus) in coastal and estuarine waters. Frontiers in Marine Science 3: 12. DOI: 10.3389/fmars.2016.00012.


Connor RC, Krützen M (2015) Male dolphin alliances in Shark Bay: changing perspectives in a thirty-year study. Animal Behaviour 103: 223-235.

King SL (2015). You talkin’ to me? Interactive playback is a powerful yet underused tool in animal communication research. Biology Letters 11: 20150403.

King SL, Schick RS, Donovan CD, Booth CG, Burgman M, Thomas L, Harwood J (2015). An Interim Framework for Assessing the Population Consequences of Disturbance. Methods in Ecology and Evolution 6(10): 1150–1158.

King SL, Janik VM (2015). Come dine with me: food associated social signalling in wild bottlenose dolphins (Tursiops truncatus)Animal Cognition 18(4):969-74.

Vollmer NL, Hayek LAC, Heithaus MR, Connor RC (2015). Further evidence of a context-specific agonistic signal in bottlenose dolphins: the influence of consortships and group size on the pop vocalization. Behavior 152: 1979-2000. 


Allen SJ (2014). From exploitation to adoration: The historical and contemporary contexts of human-cetacean interactions. In Whale-Watching: Sustainable tourism and ecological management (eds. Higham JES et al.). Cambridge University Press: Cambridge, UK. Pp. 31-47.

Allen SJ, Tyne J, Kobryn HT, Bejder L, Pollock KH, Loneragan NR (2014). Patterns of dolphin bycatch in a north-western Australian trawl fishery. PLoS ONE 9: e93178.

Brown A, Kopps AM, Allen SJ, Bejder L, Littleford-Colquhoun B, Parra GJ, Cagnazzi D, Thiele D, Palmer C, Frère C (2014). Population differentiation and hybridisation of Australian snubfin (Orcaella heinsohni) and Indo-Pacific humpback dolphins (Sousa chinensis) in north-western Australia. PLoS ONE 9: e101427.

Furuichi T, Connor R, Hashimoto C (2014). Non-conceptive sexual interactions in monkeys, apes, and dolphins. In Primates and Cetaceans (eds. J. Yamagiwa & L. Karczmarski). Springer: Japan. Pp 385–408.

Jaiteh VF, Allen SJ, Meeuwig JJ, Loneragan NR (2014). Combining in-trawl video with observer coverage improves understanding of protected and vulnerable species bycatch in trawl fisheries. Marine & Freshwater Research 65(9): 830-837.

King SL, Harley HE, & Janik VM (2014). The role of signature whistle matching in bottlenose dolphins (Tursiops truncatus). Animal Behaviour 96: 79-86

Kopps AM, Ackermann CY, Sherwin WB, Allen SJ, Bejder L, Krützen M (2014). Cultural transmission of tool use combined with habitat specializations leads to fine-scale genetic structure in bottlenose dolphins. Proceedings of the Royal Society B 281: 20133245.

Kopps AM, Krützen M, Allen SJ, Bacher K, Sherwin WB (2014). Characterizing the socially transmitted foraging tactic “sponging” by bottlenose dolphins (Tursiops sp.) in the western gulf of Shark Bay, Western Australia. Marine Mammal Science 30: 847-863.

Krützen M, Kreicker S, MacLeod CD, Learmonth J, Kopps AM, Walsham P, Allen SJ (2014). Cultural transmission of tool use by Indo-Pacific bottlenose dolphins (Tursiops sp.) provides access to a novel foraging niche. Proceedings of the Royal Society B 281: 20140374.


Allen SJ (2013). Common bottlenose dolphin Tursiops truncatus. Species account. In Field Companion to the Mammals of Australia (eds. Van Dyck S et al). New Holland Press: Sydney, Australia.

Allen SJ, Krützen M (2013). Indo-Pacific bottlenose dolphin Tursiops aduncus. Species account. In Field Companion to the Mammals of Australia (eds. Van Dyck S et al). New Holland Press: Sydney, Australia.

Jaiteh VF, Allen SJ, Meeuwig JJ, Loneragan NR (2013). Subsurface behaviour of bottlenose dolphins (Tursiops truncatus) interacting with fish trawl nets in north-western Australia. Marine Mammal Science 29(3): E266-E281.

Janik VM, King SL, Sayigh LS, Wells RS (2013). Identifying signature whistles from recordings of groups of unrestrained bottlenose dolphins (Tursiops truncatus). Marine Mammal Science 29(1):109-122.

King SL, Janik VM (2013). Bottlenose dolphins can use learned vocal labels to address each other. Proceedings of the National Academy of Sciences USA 110(32): 13216-13221

King SL, Sayigh L, Wells R, Fellner W, Janie VM (2013). Vocal copying of individually distinctive signature whistles in bottlenose dolphins. Proceedings of the Royal Society B 280: 1757.


Allen SJ, Cagnazzi DD, Hodgson AJ, Loneragan NR, Bejder L (2012). Tropical inshore dolphins of north-western Australia: Unknown quantities along a changing coastline. Pacific Conservation Biology 18(1): 56-63.

Brown A, Bejder L, Cagnazzi D, Parra G, Allen SJ (2012). The North West Cape, Western Australia: A potential hotspot for Indo-Pacific Humpback Dolphins Sousa chinensis? Pacific Conservation Biology 18(4): 240-246.

Kopps AM, Sherwin WB (2012). Modelling the emergence and stability of a vertically transmitted cultural trait in bottlenose dolphins. Animal Behaviour 84: 1347-1362.

Krzyszczyk E, Kopps AM, Bacher K, Smith H, Stephens N, Meighan NA, Mann J (2012). A report on six cases of seagrass-associated gastric impaction in bottlenose dolphins (Tursiops sp.) Marine Mammal Science doi:10.1111/j.1748-7692.2012.00579.x.

Nicholson K, Bejder L, Allen SJ, Krützen M, Pollock KH (2012). Abundance, survival and temporary emigration of bottlenose dolphins (Tursiops sp.) off Useless Loop in the western gulf of Shark Bay, Western Australia. Marine and Freshwater Research 63: 1059-1068.

Randić S, Connor R, Sherwin W, Krützen M (2012). A novel mammalian social structure in Indo-Pacific bottlenose dolphins (Tursiops sp.): complex male alliances in an open social network. Proceedings of the Royal Society B 279: 3083-3090.

Tyne J, Loneragan N, Kopps A, Allen S, Krützen M, Bejder, L (2012) Ecological characteristics contribute to sponge distribution and tool use in bottlenose dolphins Tursiops sp. Marine Ecology Progress Series 444: 143-153.


Allen SJ, Bejder L, Krützen M (2011). Why do Indo-Pacific bottlenose dolphins (Tursiops sp.) carry conch shells (Turbinella sp.) in Shark Bay, Western Australia? Marine Mammal Science 27: 449-454.

Connor RC, Watson-Capps JJ, Sherwin WB, Krützen M (2011). A new level of complexity in the male alliance networks of indian ocean bottlenose dolphins (Tursiops sp.). Biology Letters 7: 623–626.

Frère C, Mann J, Krützen M, Connor R, Bejder L, Sherwin W (2011). Nature and nurture: A step towards investigating their interactions in the wild. Communicative and Integrative Biology 4: 192-193.


Alberts S, Altman J, Bejder L, Connor R, Daher MA, Engleby L, Flaherty C, Primack D, Read A, Ridgway SH, Silk J, Spradlin T, Tyack P (2010) Memories: Amy Samuels. Marine Mammal Science 26: 501-507.

Aureli F, Schaffner C, Boesch C, Bearder S, Call J, Chapman C, Connor R, Di Fiore A, Dunbar R, Henzi S, et al. (2008). Fission-fusion dynamics. Current Anthropology 49: 627–654.

Bacher K, Allen S, Lindholm AK, Bejder L, Krützen M (2010). Genes or Culture: Are Mitochondrial Genes Associated with Tool Use in Bottlenose Dolphins (Tursiops sp.)? Behavior Genetics 40: 706-714.

Bejder L, Samuels A, Whitehead H, Finn H, Allen SJ (2009). Impact assessment research: use and misuse of habituation, sensitisation and tolerance in describing wildlife responses to anthropogenic stimuli. Marine Ecology Progress Series 395: 177-185.

Bejder L, Samuels A, Whitehead H, Gales N, Mann J, Connor R, Heithaus M, Watson-Capps J, Flaherty C, Krützen M (2006). Decline in relative abundance of bottlenose dolphins exposed to long-term disturbance. Conservation Biology 20: 1791–1798.

Connor RC (2007). Dolphin social intelligence: complex alliance relationships in bottlenose dolphins and a consideration of selective environments for extreme brain size evolution in mammals. Philosophical Transactions of the Royal Society B 362: 587-602.

Connor RC (2010). Cooperation beyond the dyad: on simple models and a complex society. Philosophical Transactions of the Royal Society B 365: 2687–2697.

Connor RC, Mann J (2006). Social cognition in the wild: Machiavellian dolphins? In Rational Animals? (eds. Hurley S, Nudds M). Oxford University Press: Oxford. Pp 329–367

Connor R, Mann J, Watson-Capps J (2006). A sex-specific affiliative contact behavior in Indian Ocean bottlenose dolphins, Tursiops sp. Ethology 112: 631–638.

Connor R, Smolker R, Bejder L (2006). Synchrony, social behaviour and alliance affiliation in Indian Ocean bottlenose dolphins, Tursiops aduncusAnimal Behaviour 72: 1371–1378.

Connor RC, Vollmer N (2009). Sexual coercion in dolphin consortships: a comparison with chimpanzees. In Sexual coercion in primates: an evolutionary perspective on male aggression against females (eds. M.N. Muller & R.W. Wrangham ). Pp 218–243. Cambridge: Harvard University Press.

Frère C, Krützen M, Mann J, Watson-Capps J, Tsai Y, Patterson E, Connor R, Bejder L,  Sherwin W (2010). Home range overlap, matrilineal and biparental kinship drive female associations in bottlenose dolphins. Animal Behaviour 80: 481–486.

Frère CH, Krützen M, Mann J, Connor RC, Bejder L, Sherwin WB (2010). Social and genetic interactions drive fitness variation in a free-living dolphin population. Proceedings of the National Academy of Sciences 107: 19949–19954.

Frère C, Krützen M, Kopps A, Ward P, Mann J, Sherwin WB (2010). Inbreeding Tolerance and Fitness Costs in Wild Bottlenose Dolphins. Proceedings of the Royal Society B 277: 2667-2673.

Gilby IC, Connor RC (2010). The role of intelligence in group hunting: are chimpanzees different from other social predators. In The mind of the chimpanzee: ecological and experimental perspectives (eds. Lonsdorf EV et al.). University of Chicago Press: Chicago. Pp 220–233.

Krützen M, Allen SJ (2008). Indo-Pacific Bottlenose Dolphin Tursiops aduncus. Species account. In The Mammals of Australia (eds van Dyck, Strahan R). New Holland Publishers: Sydney. Pp 837-838

Mann J, Sargeant BL, Watson-Capps JJ, Gibson QA, Heithaus MR, Connor RC, Patterson E (2008). Why Do Dolphins Carry Sponges? PLoS ONE 3: 1-7.

Marino L, Connor RC, Fordyce RE, et al. (2007). Cetaceans have complex brains for complex cognition. PLoS Biology 5: e139.

Nater A, Kopps AM, Krützen M (2009). New polymorphic tetranucleotide microsatellites improve scoring accuracy in bottlenose dolphins (Tursiops aduncus). Molecular Ecology Resources 9: 531-534.

Tyne JA, Loneragan NR, Krützen M, Allen SJ, Bejder L (2010). An integrated data management and video system for sampling aquatic benthos. Marine and Freshwater Research 61: 1023-1028.


Connor RC (2001). Individual foraging specializations in marine mammals: culture and ecology. Behavioral and Brain Sciences 24: 329-330.

Connor RC (2001). Social relationships in a big-brained aquatic mammal. In Model Systems in Behavioral Ecology (ed. Dugatkin LA).Princeton University Press. Pp 408-432.

Connor RC, Heithaus M, Barre L (2001) Complex social structure, alliance stability, and mating access in bottlenose dolphin ’super-alliance’. Proceedings of the Royal Society B 286: 263–267.

Connor RC, Krützen M (2003). Levels and patterns in dolphin alliance formation. In Animal Social Complexity: Intelligence, Culture, and Individualized Societies (eds. de Waal F, Tyack PL). Harvard University Press: Cambridge. Pp. 115–120.

Connor RC, Whitehead H (2005). Alliances II: Rates of encounter during resource utilization: a general model of intrasexual alliance formation in fission-fusion societies. Animal Behaviour 69: 127–132.

Gero S, Bejder L, Whitehead H, Mann J, Connor RC (2005). Behaviorally specific preferred associations in bottlenose dolphins, Tursiops sp. Canadian Journal of Zoology 83: 1566-1573.

Krützen M, Barre LM, Connor RC, Mann J, Sherwin WB (2004). O father: where art thou? – Paternity assessment in an open fission-fusion society of wild bottlenose dolphins (Tursiops sp.) in Shark Bay, Western Australia. Molecular Ecology 13: 1975-1990.

Krützen M, Barre LM, Möller LM, Simms C, Heithaus MR, Sherwin WB (2002). A biopsy system for small cetaceans: darting success and wound healing in Tursiops sp. Marine Mammal Science 18: 863-878.

Krützen M, Mann J, Heithaus M, Connor R, Bejder L, Sherwin WB (2005). Cultural transmission of tool use in bottlenose dolphins. Proceedings of the National Academy of Sciences USA 105: 8939-8943.

Krützen M, Sherwin WB, Berggren P, Gales NJ (2004). Population structure in an inshore cetacean revealed by microsatellite and mtDNA analysis: bottlenose dolphins (Tursiops sp.) in Shark Bay, Western Australia. Marine Mammal Science 20: 28-47.

Krützen M, Sherwin WB, Connor RC, Barré LM, Van de Casteele T, Mann J, Brooks R (2003). Contrasting relatedness patterns in bottlenose dolphins (Tursiops sp.) with different alliance strategies. Proceedings of the Royal Society of London B 270: 497–502.

Krützen M, Valsecchi E, Connor R, Sherwin WB (2001). Characterization of microsatellite loci in Tursiops aduncusMolecular Ecology Notes 1: 170–172.

Sargeant B, Mann J, Berggren P, Krützen M (2005). Specialization and development of beach hunting, a rare foraging behavior, by wild bottlenose dolphins (Tursiops sp.). Canadian Journal of Zoology 83: 1400-1410.

Scott EM, Mann J, Watson JJ, Sergeant BL, Connor RC (2005). Aggression in bottlenose dolphins: Evidence for sexual coercion, male-male competition, and female tolerance through analysis of tooth-rake marks and behaviour. Behaviour 142: 21-44.

Whitehead H, Connor RC (2005). Alliances I: How large should alliances be? Animal Behaviour 69: 117-126.


Connor RC (2000). Group Living in Whales and Dolphins. In Cetacean Societies: Field studies of whales and dolphins (eds. Mann J, Connor R, Tyack P, Whitehead H). University of Chicago Press: Chicago. Pp. 199-218.

Connor RC, Heithaus MR, Berggren P, Miksis JL (2000). ‘Kerplunking’: Surface fluke-splashes during shallow water bottom foraging by bottlenose dolphins. Marine Mammal Science 16: 646-653.

Connor RC, Read A, Wrangham RW (2000). Male reproductive strategies and social bonds. In Cetacean Societies: Field studies of whales and dolphins (eds. Mann J, Connor R, Tyack P, Whitehead H). University of Chicago Press: Chicago. Pp. 247-269.

Connor R, Wells R, Mann J, Read A (2000). The bottlenose dolphin. In Cetacean Societies: Field studies of whales and dolphins (eds. Mann J, Connor R, Tyack P, Whitehead H). University of Chicago Press: Chicago. Pp. 91–125.

Connor RC, Heithaus M, Barre L (1999). Super-alliance of bottlenose dolphins. Nature 371: 571–572.

Connor RC, Mann J, Tyack PL, Whitehead H (1998). Social evolution in toothed whales. Trends in Ecology and Evolution 13: 228–232.

Connor RC, Heithaus MR (1996). Approach by great white shark elicits flight response in bottlenose dolphins. Marine Mammal Science 12: 602-606.

Connor R, Richards A, Smolker R, Mann, J (1996). Patterns of female attractiveness in indian ocean bottlenose dolphins. Behaviour 133: 37–69.

Connor R, Smolker R (1996). ‘Pop’ goes the dolphin: A vocalization male bottlenose dolphins produce during consortships. Behaviour 133: 643–662.

Mann J, Connor R, Barre L, Heithaus M (2000). Female reproductive success in bottlenose dolphins (Tursiops sp.): Life history, habitat, provisioning, and group-size effects. Behavioral Ecology 11: 210–219.

Smolker R, Pepper JW (1999). Whistle convergence among allied male bottlenose dolphins (Delphinidae, Tursiops sp.). Ethology 105: 595–617.

Smolker R, Richards A, Connor R, Mann J, Berggren P (1997). Sponge Carrying by Dolphins (Delphinidae, Tursiops sp.): A Foraging Specialization Involving Tool Use? Ethology 103: 454-465.


Connor R, Smolker R (1995). Seasonal changes in the stability of male-male bonds in Indian Ocean bottlenose dolphins (Tursiops sp.). Aquatic Mammals 21: 213–216.

Connor RC, Smolker RA, Richards AF (1992). Dolphin alliances and coalitions. In Coalitions and Alliances in Animals and Humans (eds. Harcourt A, de Waal F). Oxford University Press: Oxford. Pp. 415–443.

Connor RC, Smolker RA, Richards AF (1992). Two levels of alliance formation among male bottlenose dolphins (Tursiops sp). Proceedings of the National Academy of Sciences USA 89: 987–990.

Mann J, Smolker RA, Smuts BB (1995). Responses to calf entanglement in free-ranging bottlenose dolphins. Marine Mammal Science 11: 100–106.

Smolker R, Mann J, Smuts B (1993). Use of signature whistles during separations and reunions by wild bottlenose dolphin mothers and infants. Behavioral Ecology & Sociobiology 33: 393–402.

Smolker RA, Richards AF, Connor RC, Pepper J (1992). Sex differences in patterns of association among Indian Ocean bottlenose dolphins. Behaviour 123: 38–69.


Connor RC, Smolker RA (1990). Quantitative description of a rare behavioral event: a bottlenose dolphins behavior toward her deceased offspring. In The Bottlenose Dolphin(eds. Leatherwood S, Reeves RR). Academic Press: New York. Pp. 355-360.

Connor RC, Smolker RA (1985). Habituated dolphins (Tursiops sp.) in Western Australia. Journal of Mammalogy 66: 398-400.

Smolker R, Richards AF (1988). Loud sounds during feeding in Indian Ocean bottlenose dolphins. In Animal Sonar: processes and performance (eds. Nachtigall P, Moore PWB). Plenum Press: New York. Pp. 703-706.